Matsumurella Makino

Head. Head subequal to or wider than pronotum. Crown not produced, apex not longer than 1.5x length next to eye, flat or convex. Discal portion of crown radially or longitudinally striate. Transition of crown to face rounded. Texture of anterior margin of head shagreen. Frontoclypeus moderately narrow, median length 1.25x to 2x width between antennal pits, not widening abruptly dorsad of antennal pits, texture shagreen. Clypellus parallel sided or tapering apically, relatively flat. Lorum subequal to or wider than clypellus near base. Antennal bases near middle or posteroventral (lower) corners of eyes. Antennae short, less than 1.5x width of head. Genae incised laterally. Fine erect seta on gena close to lateral frontal suture. Maxillary sensillum closely adjacent to lorum. Lorum closely adjacent to lower genal margin. Antennal ledges weakly developed (carinate or weakly carinate). Mesial margin of eye entire. Eye less than .5x width of crown. Ocelli medium sized to large, close to eyes, less than 1/3 distant from eye to crown apex, on anterior margin, partially visible in dorsal view.

Thorax. Pronotum lateral margin carinate, lateral margin less than 1/2 basal width of eye or lateral margin longer than 1/2 basal width of eye, with tranverse striations.

Wings. Forewing rounded apically, macropterous, transparent, appendix restricted to anal margin, with 3 anteapical cells, outer anteapical cell rounded or truncate apically, not particularly small, with 1 or more extra straight costal veins, not reflexed, extra costal veins arising from outer anteapical cell and/or basad of R1, without extra crossveins on corium or with a few extra crossveins on corium or reticulate, with many extra crossveins on corium, without extra crossveins in brachial cell or with extra crossveins in brachial cell, without false veins, with one crossvein between A1 and claval suture or with 2 or more crossveins between A1 and claval suture, anal veins not strongly curved, meeting commisural margin obliquely, claval cells without definitely delimited transparent or pale opaque spots, A1-A2 crossvein present, inner anteapical cell open or inner anteapical cell closed. Hindwing fully developed. Hind wing complete. Hind wing veins R4+5 and M1+2 separated by a crossvein or touching for a very short distance, M3+4 and CuA separated by a crossvein. Hind wing with 2 branches.

Legs. Profemur with AM1 seta only, AM 1 seta near mid-height of femur. Profemur intercalary row setae thin. Profemur number of intercalary row setae: 7-15 . Profemur row AV with short, stout macrosetae. Profemur number of row AV setae 20-35 . Profemur AV1 seta poorly differentiated from intercalary row setae. Protibia row AD with 2 to 4 macrosetae or row AD with 5 or more macrosetae. Protibia row PD number of macrosetae: 4-5 . Mesotrochanter with one stout apical seta. Mesofemur row AV with several to numerous short stout setae. Metafemur apex penultimate set of macrosetae with 2 setae, antepenultimate set of macrosetae with 1 seta, without extra macrosetae proximal of typical 2+2+1. Metatibia proximal seta of PD distad of proximal seta of AD or equidistant from tibia base, row AD intercalary setae simple, without basal stout setae on dorsal surface, row PD macrosetae subequal in length to AD macrosetae, row PD macrosetae alternating short and long, basal dorsal width subequal to width near apex, pecten with median seta much longer than adjacent setae. Metatarsomere I not expanded apically, long, slender, plantar setae all simple. Metatarsomere I pecten: number of platellae 5-6 . Metatarsomere I pecten inner apical seta platelliform.

Male Genitalia. Valve articulated with pygofer and subgenital plates, with short point of articulation with pygofer. Pygofer dorsoapical margin incised nearly to base, macrosetae absent or reduced (<= two rows) or macrosetae well differentiated into several rows, without dense tufts of thick or fine setae, without ventral tooth, spine, or process, with dorsal tooth, spine, or process, posterior margin without teeth or serrations. Segment X sclerotized laterally and dorsally, not particularly long or broad. Subgenital plates triangular, longer than width, outer margin convex, apex pointed or apex rounded or medial margin notched or excavated subapically, with two lateral rows of macrosetae, or with some irregularly arranged macrosetae near lateral margin, without or with few long fine setae. Style preapical lobe reduced or absent or preapical lobe short, rounded or pointed, apophysis shorter than rest of style, apophysis straight or apophysis smoothly curved, apophysis apex rounded, apophysis inner (posterior) margin without teeth, apophysis smooth. Aedeagus without processes arising from base, without medial processes, with one pair of apical processes, depressed, flagellum arising from ventral part of socle, symmetrical, moderate to long, gonoduct not sclerotized basad of aedeagal base, gonopore subapical, on dorsal side, with a single shaft and gonopore, preatrium absent or very short. Phragma membranous. Connective anterior arms somewhat divergent, (Y- or U-shaped), stem present, shorter than anterior arms, articulated with aedeagus, without processes.