@article {Braeuchler:2010p2765,
	title = {Molecular phylogeny of Menthinae (Lamiaceae, Nepetoideae, Mentheae) - Taxonomy, biogeography and conflicts},
	journal = {Molecular Phylogenetics and Evolution},
	volume = {55},
	number = {2},
	year = {2010},
	month = {Jan},
	pages = {501{\textendash}523},
	abstract = {Although the subfamily Nepetoideae (Lamiaceae) is considered to be monophyletic, relationships between tribes, subtribes and genera within the subfamily are poorly understood as complex and possibly homoplasious morphological characters make taxa difficult to delimit. DNA sequence data from three regions (chloroplast: trnK intron; trnL-F; nuclear: ITS) in total including 278 accessions, representing 38 out of 40 genera of subtribe Menthinae and 11 outgroup genera, were used to reconstruct the evolutionary history, test previous hypotheses of classification, explain biogeographic patterns and elucidate character evolution. Using maximum parsimony (MP) and Bayesian analysis phylogenetic reconstructions based on nuclear and chloroplast sequence data were incongruent, consequently the data were analyzed separately.Both nuclear and chloroplast datasets provide strong support for three major lineages: the "Satureja", "Micromeria" and "Clinopodium" group. The first contains members of Satureja and Gontscharovia. In the second lineage Micromeria s.str. and Origanum were resolved as monophyletic, Pentapleura and Zataria indicated as sister groups. Thymbra includes two species of Satureja turning the latter genus polyphyletic. Thymus is revealed as paraphyletic with respect to Argantoniella and Saccocalyx in both and Origanum in the plastid dataset only. In the third lineage, the Clinopodium-group, branching pattern is highly incongruent among datasets and possibly influenced by recent and ancient hybridization, chloroplast capture and incomplete lineage sorting. However, identical terminal groups are inferred in both analyses. A Madagascan lineage of "Micromeria", sister to the recently described South African Killickia, is suggested to represent a new genus. The Himalayan Clinopodium nepalense group and the tropical African C. abyssinicum alliance are monophyletic but indicated in different positions. Both groups appear in the ITS phylogeny in a clade with Cyclotrichium and Mentha, relationships not suggested previously. The enigmatic Micromeria cymuligera is close to Mentha and possibly is a representative of this genus. Species of Acinos, now regarded as part of Clinopodium, are mixed up with species of Ziziphora, questioning either the inclusion of Acinos in Clinopodium or generic distinctness of Ziziphora. All data sets suggest a monophyly of the New World taxa and argue for long distance dispersal from the Old World, rather than a vicariance explanation. Bystropogon marks the split up between the two lineages. Inclusion of 22 genera intermixed with Clinopodium spp. in the New World clade provides evidence that the current circumscription of the genus is highly unnatural. Low sequence divergence resulting in low phylogenetic resolution especially at the base of the clade indicate a rapid radiation accompanied by considerable ecological diversification and speciation. (C) 2010 Elsevier Inc. All rights reserved.},
	keywords = {Chloroplast Dna, Generic delimitations, Genus Bystropogon Lamiaceae, Island Colonization, Lamiaceae, Menthinae, Micromeria Lamiaceae, Minthostachys Labiatae, Molecular phylogeny, North-America, Nuclear Ribosomal Dna, Parsimony Analysis, Salvia Lamiaceae, Satureja, Scrub Mints Lamiaceae},
	doi = {10.1016/j.ympev.2010.01.016},
	url = {http://www.sciencedirect.com/science?_ob=ArticleURL\&_udi=B6WNH-4YC2XFX-1\&_user=2139813\&_coverDate=05\%252F31\%252F2010\&_rdoc=1\&_fmt=high\&_orig=gateway\&_origin=gateway\&_sort=d\&_docanchor=\&view=c\&_acct=C000054276\&_version=1\&_urlVersion=0\&_userid=2139813\&md5=d},
	author = {Christian Braeuchler and Harald Meimberg and Guenther Heubl}
}
@article {Scheen:2010p3190,
	title = {MOLECULAR PHYLOGENETICS, CHARACTER EVOLUTION, AND SUPRAGENERIC CLASSIFICATION OF LAMIOIDEAE (LAMIACEAE)},
	journal = {Annals of the Missouri Botanical Garden},
	volume = {97},
	number = {2},
	year = {2010},
	month = {Jan},
	pages = {191{\textendash}217},
	abstract = {This paper presents a phylogenetic analysis of Lamiaceae subfam. Lamioideae (including subfamily Pogostemonoideae) based on sequences of the trnL intron, trnL-trnF intergenic spacer, and rps16 intron of the plastid genuine. It is the first analysis that includes all major lamioid and pogostemonoid genera. Monophyly of Lamioideae s.l. (i.e., including Pogostemonoideae) is strongly supported, with Cymaria Benth. as its sister group, and Pogostemonoideae, which sometimes has been recognized as a subfamily, is subsumed in Lamioideae. On the basis of the phylogenetic hypothesis, Lamioideae is divided into nine tribes. Three new tribes are established: Gomphostemmateae Seheen {\&} Lindqvist, Phlomideae Mathiesen, and Leucadeae Scheen {\&} Ryding. The other six tribes are: Pogostemoneae Briq., Synandreae Raf., Staehydeae Dumort., Leonureae Dumort., Limieae Coss. {\&} Germ., and Marrubieae Vis. The genus Betonica L. is reestablished. The results also strongly suggest that the genera Stachys L., Sideritis L., Ballota and Leucas R. Br. are polyphyletic or paraphyletic. The results were used to examine evolution of non-molecular characters.},
	keywords = {Character Evolution, Chloroplast Dna, Classification, Eremostachys Bunge, Hawaiian Endemic Mints, Lamiaceae, Lamioideae, Molecular Phylogeneties, Morphology, Pericarp Structure, Phlomis L, Pogostemonoideae, Rps16 Intron, Sequence Data, Subtribe Melittidinae Labiatae, Systematic Implications, Verbenaceae},
	doi = {10.3417/2007174},
	url = {http://www.bioone.org/doi/abs/10.3417/2007174},
	author = {Anne-Cathrine Scheen and Mika Bendiksby and Olof Ryding and Cecilie Mathiesen and Victor A Albert and Charlotte Lindqvist}
}
