@article {Bendiksby:2011p3198,
	title = {Molecular phylogeny and taxonomy of the genus Lamium L. (Lamiaceae): Disentangling origins of presumed allotetraploids},
	journal = {Taxon},
	volume = {60},
	number = {4},
	year = {2011},
	month = {Jan},
	pages = {986{\textendash}1000},
	abstract = {This is the first comprehensive molecular investigation of the genus Lamium L. We have addressed phylogenetic relationships and presumed allopolyploid speciation by use of nuclear (NRPA2, 5S-NTS) and chloroplast (matK, psbA-trnH, rps16, trnL, trnL-F, trnS-G) DNA sequence data. Nuclear and chloroplast data were incongruent, and nuclear data showed better correlation with morphology. Bayesian and parsimony phylogenetic results show that (I) Lamium galeobdolon is sister to all remaining Lamium species; (2) Wiedemannia is nested within Lamium; (3) L. amplexicaule is polyphyletic; (4) most tetraploids are of hybrid origin; (5) L. amplexicaule var. orientale is allotetraploid; and (6) Mennema{\textquoteright}s (1989) infrageneric classification is not corroborated by molecular data. Based on the molecular results, and taking morphology into account, we suggest resurrection of two species: L. aleppicum and L. paczoskianum.},
	keywords = {5S-Nts, Allopolyploidy, Asteraceae, Chloroplast Dna, Classification, Complex, Cpdna, Dna-Sequences, Incongruence, Lamioideae, Lamium, Molecular Phylogenetics, Nrpa2, Reconstruction, Reticulate Evolution, Silene Caryophyllaceae, Speciation},
	url = {http://apps.webofknowledge.com/InboundService.do?SID=2B\%2540kNem2NICHnHfl3PI\&product=WOS\&UT=000294507000004\&SrcApp=Papers\&DestFail=http\%253A\%252F\%252Faccess.isiproducts.com\%252Fcustom_images\%252Fwok5_failed_auth.html\&Init=Yes\&action=retrieve\&SrcAuth=meken},
	author = {Mika Bendiksby and Anne K Brysting and Lisbeth Thorbek and Galina Gussarova and Olof Ryding}
}
@article {Ryding:1995p1138,
	title = {Pericarp structure and phylogeny of the Lamiaceae-Verbenaceae-complex},
	journal = {Plant Systematics and Evolution},
	volume = {198},
	number = {1-2},
	year = {1995},
	month = {Jan},
	pages = {101{\textendash}141},
	abstract = {Pericarp structure was investigated in 158 species of the families Lamiaceae and Verbenaceae. Data from 221 out of 262 genera of Lamiaceae s.l. and a few of Verbenaceae s.str. were collected in a table. A cladistic analysis was performed on the basis of pericarp characters only. The abandonment of subfam. Pogostemonoideae as a taxonomic unit is considered. Examples of groups given additional support by similarities in pericarp characters are: (1) the gynobasic-styled labiates (subfamilies Pogostemonoideae, Lamioi deae, Nepetoideae); (2) a Lamioideae-Pogostemonoideae-group; (3) Nepetoideae; (4) a Westringia-Hemigenia-Hemiandra-Microcorys group (in subfam. Chloranthoideae); (5) a Lepechinia-Chaunostoma-group (in Nepetoideae); (6) a Prunella-Cleonia-group (in Nepetoideae).},
	keywords = {Anatomy, Chloanthoideae, Classification, Labiatae, Lamiaceae, Lamioideae, Morphology, Nepetoideae, Pericarp, Phylogeny, Pogostemonoideae, Rbcl, Scutellarioideae, Sequences, Verbenaceae},
	url = {http://apps.isiknowledge.com/InboundService.do?product=WOS\&action=retrieve\&SrcApp=Papers\&UT=A1995TN62000006\&SID=2AFL5dnj5MCc8JPedL5\&SrcAuth=mekentosj\&mode=FullRecord\&customersID=mekentosj\&DestFail=http\%253A\%252F\%252Faccess.isiproducts.com\%252Fcustom_image},
	author = {O Ryding}
}
@article {Bendiksby:2011p3196,
	title = {An updated phylogeny and classification of Lamiaceae subfamily Lamioideae},
	journal = {Taxon},
	volume = {60},
	number = {2},
	year = {2011},
	month = {Jan},
	pages = {471{\textendash}484},
	abstract = {Lamioideae comprise the second-largest subfamily in Lamiaceae. Although considerable progress has recently been made in Lamioideae phylogenetics, the subfamily remains one of the most poorly investigated subfamilies in Lamiaceae. Here we present a taxonomic update of the subfamily based on earlier published data as well as 71 new DNA extracts from relevant in- and outgroup taxa, and DNA sequence data from four chloroplast regions (matK, rps16, trnL intron and trnL-F spacer). The phylogenetic positions of 10 out of 13 previously unplaced small or monotypic Asian lamioid genera and 37 additional lamioid species have been identified, and the classification is updated accordingly. Results from parsimony and Bayesian phylogenetic methods corroborate earlier results, but phylogenetic resolution as well as overall branch support are improved. All newly added genera are assigned to earlier established tribes or the new tribe Paraphlomideae Bendiksby, which includes Ajugoides, Matsumurella and Paraphlomis. Acanthoprasium is resurrected as a genus. Transfer of species is proposed to accommodate the monophyly of two genera (Lamium, Otostegia), whereas ten genera remain non-monophyletic (Ballota s.str., Lagopsis, Leonotis, Leonurus, Leucas, Microtoena, Phlomoides, Sideritis, Stachys, Thuspeinanta). Eriophyton and Stachyopsis have been included in Lamieae, Hypogomphia in Stachydeae, and Loxocalyx in Leonureae. Betonica, Colquhounia, Galeopsis, and Roylea remain unclassified at the tribal level. Lamium chinense and three East Asian Galeobdolon species are transferred to Matsumurella. Sulaimania and four Otostegia species are transferred to Moluccella. Alajja and three Lamium species are transferred to Eriophyton. In total, 14 new combinations are made, one at the rank of subgenus and 13 at the rank of species.},
	keywords = {Acanthoprasium, Chloroplast Sequence Datasets, Classification, Dna, Eriophyton, Evolution, Hawaiian Endemic Mints, Labiatae, Lamiaceae, Lamioideae, Matsumurella, Molecular Phylogenetics, Moluccella, Nuclear, Origin, Paraphlomideae, Pericarp Structure, Sideritis Lamiaceae},
	url = {http://apps.webofknowledge.com/InboundService.do?SID=2B\%2540kNem2NICHnHfl3PI\&product=WOS\&UT=000289587100015\&SrcApp=Papers\&DestFail=http\%253A\%252F\%252Faccess.isiproducts.com\%252Fcustom_images\%252Fwok5_failed_auth.html\&Init=Yes\&action=retrieve\&SrcAuth=meken},
	author = {Mika Bendiksby and Lisbeth Thorbek and Anne-Cathrine Scheen and Charlotte Lindqvist and Olof Ryding}
}
@article {Ryding:2007p1121,
	title = {Amount of calyx fibres in Lamiaceae, relation to calyx structure, phylogeny and ecology},
	journal = {Plant Systematics and Evolution},
	volume = {268},
	number = {1-4},
	year = {2007},
	month = {Jan},
	pages = {45{\textendash}58},
	abstract = {Calyces of 306 species representing 181 out of the 236 genera of Lamiaceae have been studied with particular emphasis on the amount of fibres and similar xylem cells. A characteristic calyx tissue called "mesophyll fibres", reported by earlier authors, is found to belong to the xylem. Most species of the subfamilies Lamioideae and Scutellarioideae differ from most other labiates, and particularly from Nepetoideae, in having much larger amounts of fibres and similar xylem cells in the calyx tube. This result supports cladistic hypotheses based on cpDNA showing that Lamioideae and Scutellarioideae are closely related and remotely related to Nepetoideae. The new data also contribute to the knowledge about the phylogeny within Lamioideae. The amount of fibres and similar cells also seems to be positively correlated with calyx size, calyx width, a ballistic dispersal mechanism in Scutellaria, and the aridity of the habitat. Possible adaptive significances of these correlations are discussed.},
	keywords = {Calyx, Fibres, Labiatae, Lamiaceae, Lamioideae, Phylogeny, Scutellarioideae, Sequences},
	doi = {10.1007/s00606-007-0537-y},
	url = {http://www.springerlink.com/content/51h118122296762q/},
	author = {O Ryding}
}
@article {Scheen:2010p3190,
	title = {MOLECULAR PHYLOGENETICS, CHARACTER EVOLUTION, AND SUPRAGENERIC CLASSIFICATION OF LAMIOIDEAE (LAMIACEAE)},
	journal = {Annals of the Missouri Botanical Garden},
	volume = {97},
	number = {2},
	year = {2010},
	month = {Jan},
	pages = {191{\textendash}217},
	abstract = {This paper presents a phylogenetic analysis of Lamiaceae subfam. Lamioideae (including subfamily Pogostemonoideae) based on sequences of the trnL intron, trnL-trnF intergenic spacer, and rps16 intron of the plastid genuine. It is the first analysis that includes all major lamioid and pogostemonoid genera. Monophyly of Lamioideae s.l. (i.e., including Pogostemonoideae) is strongly supported, with Cymaria Benth. as its sister group, and Pogostemonoideae, which sometimes has been recognized as a subfamily, is subsumed in Lamioideae. On the basis of the phylogenetic hypothesis, Lamioideae is divided into nine tribes. Three new tribes are established: Gomphostemmateae Seheen {\&} Lindqvist, Phlomideae Mathiesen, and Leucadeae Scheen {\&} Ryding. The other six tribes are: Pogostemoneae Briq., Synandreae Raf., Staehydeae Dumort., Leonureae Dumort., Limieae Coss. {\&} Germ., and Marrubieae Vis. The genus Betonica L. is reestablished. The results also strongly suggest that the genera Stachys L., Sideritis L., Ballota and Leucas R. Br. are polyphyletic or paraphyletic. The results were used to examine evolution of non-molecular characters.},
	keywords = {Character Evolution, Chloroplast Dna, Classification, Eremostachys Bunge, Hawaiian Endemic Mints, Lamiaceae, Lamioideae, Molecular Phylogeneties, Morphology, Pericarp Structure, Phlomis L, Pogostemonoideae, Rps16 Intron, Sequence Data, Subtribe Melittidinae Labiatae, Systematic Implications, Verbenaceae},
	doi = {10.3417/2007174},
	url = {http://www.bioone.org/doi/abs/10.3417/2007174},
	author = {Anne-Cathrine Scheen and Mika Bendiksby and Olof Ryding and Cecilie Mathiesen and Victor A Albert and Charlotte Lindqvist}
}
@article {Wink:1996p1139,
	title = {Phylogenetic relationships between some members of the subfamily Lamioideae (family Labiatae) inferred from nucleotide sequences of the rbcL gene},
	journal = {Bot Acta},
	volume = {109},
	number = {2},
	year = {1996},
	month = {Jan},
	pages = {139{\textendash}148},
	abstract = {Total DNA was extracted from 25 species (8 general of the subfamily Lamioideae sensu Erdtman (family Labiatae). Using rbcl-specific primers, the rbcL gene was amplified by polymerase chain reaction (PCR) and 1420 bp were sequenced directly, or after cloning. RbcL sequences were evaluated with character-state (maximum parsimony; PAUP) and distance methods (neighbour-joining; MEGA). In agreement with classical systematics all taxa studied cluster outside the Nepetoideae. The following clades are apparent: I - Westringia/Prostanthera, II - Ajuga/Teucrium and III - Scutellaria, Stachys, Lamium and Marrubium. Lamium album, L. amplexicaule and L. garganicum appear as a sister clade to Marrubium. Both share a common ancestor with a second Lamium clade (including L. purpureum, L. maculatum and L. galeobdolon). The phylogenetic relationships are discussed with regard to the systematic classification of the subfamilies Lamioideae and Nepetoideae (sensu Erdtman).},
	keywords = {Asteridae, Labiatae, Lamioideae, Molecular phylogeny, Nucleotide Sequences, Plants, Pollen Morphology, Rbcl, Trees},
	url = {http://access.isiproducts.com/custom_images/wok_failed_auth.html},
	author = {N Wink and M Kaufmann}
}
